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高坤山老师介绍
个人简历：
高坤山 男，1958年9月出生, 教授，博士生导师
主要研究方向：藻类生理生态。1982年毕业于中国海洋大学水产学院， 83年10月作为教育部出国研究生留学日本, 86年3月获东京水产大学硕士学位，89年3月获京都大学农学博士学位。1989年4月至92年7月，任日本关西综合环境中心客座研究员；1992年8月至94年12月，夏威夷大学博士后。1995年1月至1996年12月，汕头大学副教授。1997年入选中国科学院“百人计划”研究员、博士生导师，汕头大学教授，海洋生物研究所所长。
1996年获国家杰出青年科学基金资助，已完成国家自然科学基金重点项目“大气CO2浓度升高与水生生物的关系”、面上项目及国家海洋“863”项目各一项。 目前承担国家自然科学基金重点项目“海洋植物对阳光紫外辐射响应机制的研究”（2005.1-2008.12）。
在20多年的研究历程中，在国内外刊物上表论文90多篇，其中在国际主流刊物发表论文65篇（SCI收录51篇）。多数被国际同行学者在研究论文、综述（reviews)及专著中反复引用（正面引用）。
论文与专著中多次引用。现任〖Journal of Applied Phycology〗（Kluwer Academic Publisher, SCI收录） 唯一中国籍编委 (1998－), 《水生生物学报》、《海洋科学》编委， 国际藻类学会与美国藻类学会终身会员，日本、欧洲藻类学会会员，中国藻类学会理事。国际应用藻类学会执行委员（executive committee, 2005-)。
当前研究重点：
1)海洋生态与光生物学；
2)藻类环境生理,全球环境变化与藻类关系的研究；
3)应用藻类学,经济藻类养殖或培养的生理学基础；
4)藻类光合固碳机理,微藻与海藻类无机碳利用与浓缩机制等。
近年主要论文：
1. Gao K, Wu Y, Li G, Wu H, Villafańe V and Helbling E. 2007. Solar UV-radiation drives CO2-
fixation in marine phytoplankton: A double-edged sword. Plant Physiology. 144:54-59. (IF:6.114)
2. Villafańe V, Gao K, Li P, Li G and Helbling E. 2007. Vertical mixing within the epilimnion
modulates UVR induced photoinhibition in tropical freshwater phytoplankton from southern
China. Freshwater Biology.52:1260-1270. (IF:2.797)
3. Gao K and Ye C. 2007. Photosynthetic insensitivity of Nostoc flagelliforme to solar UV radiation
while rehydrated or desiccated. Journal of Phycology. 43(4):628-635 (IF:2.502)
4. Gao K, Li G, Helbling E and Villafańe V. 2007. Variability of UVR effects on photosynthesis of
summer phytoplankton assemblages from a tropical coastal area of the South China Sea.
Photochemistry and Photobiology. 83:802-809. (IF:2.147)
5. Jiang H, Gao K and Helbling E. 2007. Effects of solar UV radiation on germination of conchospores
and morphogenesis of sporelings inPorphyra haitanensis (Rhodophyta). Marine Biology. 151:1751-
1759. (IF:1.754)
6. Gao K, Guan W and Helbling E. 2007. Effects of solar ultraviolet radiation on photosynthesis of the
marine red tide alga Heterosigma akashiwo (Raphidophyceae). Journal of Photochemistry and
Photobiology B: Biology. 86:140-148. (IF:1.597)
7. Zou D, Gao K, Xia J, Xu Z, Zhang X and Liu S. 2007. responses of dark respiration in the light to
desiccation and temperature in the intertidal macroalga, Ulva lactuca (Chorophyta) during emersion.
Phycologia. 46(3):139-146. (IF:1.271)
8. 徐军田, 高坤山. 2007. 阳光紫外辐射对绿藻石莼光化学效率的影响. 海洋学报. 29(1):129-134.
9. 关万春, 高坤山. 2007. 阳光紫外辐射对两种赤潮藻类光化学效率的影响. 水生生物学报.
31(4):153-158.
10. Zou D, Gao K and Ruan Z. 2006. Seasonal pattern of reproduction of Hizikia fusiformis
(Sargassaceae, Phaeophyta) from Nanao Island, Shantou,China. Journal of Applied Phycology. 18:
195-201. (IF:0.992)
11. Helbling E, Gao K, Ai H, Ma Z and Villafańe V. 2006.Differential responses of Nostoc sphaeroides
and Arthrospira platensis to solar ultraviolet radiation exposure. Journal of Applied Phycology. 18
(1):57-66. (IF:0.992)
12. Ma Z, Gao K and Watanabe T. 2006. Effects of dissolved organic matter on the growth and
pigments synthesis of Spirulina platensis (Arthrospira)*. Progress in Natural Science. 16(1):
50-54. (IF:0.444)
13. Wu H, Gao K, Villafańe V, Watanabe T and Helbling E. 2005. Effect of solar UV radiation on
morphology and photosynthesis of filamentous cyanobacterium Arthrospira platensis. Applied and
Environmental Microbiology. 71(9):5004-5013. (IF:3.818)
14. Villafańe V, Gao K and Helbling E. 2005. Short-and long-term effects of solar ultraviolet radiation
on the red algae Porphyridium cruentum (S. F. Gray) N?geli. Photochemical & Photobiological
Sciences. 4(4):376-382. (IF:2.117)
15. Xia J and Gao K. 2005. Impacts of elevated CO2 concentration on biochemical composition,
carbonic anhydrase, and nitrate reductase activity of freshwater green algae. Journal of Integrative
Plant Biology. 47(6):668-675. (IF:0.432)
16. Zou D and Gao K. 2005. Regulation of gamete release in the economic brown seaweed Hizikia
fusiforme (Phaeophyta). Biotechnology Letters. 27:915-918. (IF:1.108)
17. Zou D and Gao K. 2005. Photosynthetic characteristics of the economic brown seaweed Hizikia
fusiforme (Sargassaceae, Phaeophyta), with special reference to its“leaf” and receptacle.
Journal of Applied Phycology. 17:255-259. (IF:0.992)
18. Wu H, Gao K, Ma Z and Watanabe T. 2005. Effects of solar ultraviolet radiation on biomass
production and pigment contents of Spirulina platensis in commercial operations under sunny and
cloudy weather conditions. Fisheries Science. 71(2):454-456. (IF:0.661)
19. Zou D and Gao K. 2005. Ecophysiological characteristics of four intertidal marine macroalgae
during emersion along Shantou coast of China, with a special reference to the relationship of
photosynthesis and CO2. Acta Oceanologica Sinica. 24(3):105-113. (IF:0.507)
20. Ji Y, Gao K and Tanaka J. 2005. Photosynthetic recovery of desiccated intertidal seaweeds after
rehydration. Progress in Natural Science. 15(8):689-693. (IF:0.444)
21. Wu H, Gao K and Watanabe T. 2005. Effects of solar ultraviolet radiation on the photochemical
efficiency, photosynthetic pigments and biomass production of Spirulina platensis. Progress in
Natural Science. 15(5): 477-480. (IF:0.444)
22. 吴红艳, 高坤山, 渡辺辉夫. 2005. 静止和充气培养条件下短期紫外辐射对钝顶螺旋藻光化
学效率的影响. 水生生物学报.29 (6): 674-677.
23. Gao K and Ai H. 2004. Relationship of growth and photosynthesis with colony size in an edible
cyanobacterium, Ge-Xian-Mi Nostoc (Cyanophyceae). Journal of Phycology. 40(3):523-526.
(IF:2.502)
24. Jiang H and Gao K. 2004. Effects of lowering temperature during culture on the production of
polyunsaturated fatty acids in the marine diatom Phaeodactylum tricornutum (Bacillariophyceae).
Journal of Phycology. 40(4):651-654. (IF:2.502)
25. Chen X and Gao K. 2004. Photosynthetic utilisation of inorganic carbon and its regulation in the
marine diatom Skeletonema costatum. Functional Plant Biology. 31(4):1027-1033. (IF:2.496)
26. Chen X and Gao K. 2004. Characterization of diurnal photosynthetic rhythms in the marine diatom
Skeletonema costatum grown in synchronous culture under ambient and elevated CO2. Functional
Plant Biology. 31(4):399-404. (IF:2.496)
27. Li Y and Gao K. 2004. Photosynthetic physiology and growth as a function of colony size in the
cyanobacterium Nostoc sphaeroides. European Journal of Phycology. 39(1):9-15. (IF:2.064)
28. Qiu B, Zhang A, Liu Z and Gao K. 2004. Studies on the photosynthesis of the terrestrial
cyanobacterium Nostoc flagelliforme subjected to desiccation and subsequent rehydration.
Phycologia. 43(5):521-528. (IF:1.271)
29. Ye C and Gao K. 2004. Photosynthetic response to salt of aquatic-living colonies of the terrestrial
cyanobacterium Nostoc flagelliforme. Journal of Applied Phycology. 16(6):477-481. (IF:0.992)
30. Gao K and Ji Y. 2004. Quantitative evaluation of wind effect during emersion on Porhpyra
haitanensis (Rhodophyta), a farmed species in southern china. Fisheries science. 70(4):710-712.
(IF:0.661)
31. Zou D and Gao K. 2004. Comparative mechanisms of photosynthetic carbon acquisition in Hizikia
fusiforme under submersed and emersed conditions. Acta Botanica Sinica. 46(10):1178-1185.
(IF:0.432)
32. Zou D and Gao K. 2004. Exogenous carbon acquisition of photosynthesis in Porphyra haitanensis
(Bangiales, Rhodophyta) under emersed state. Progress in Natural Science. 14(2):138-144.
(IF:0.444)
33. Chen X and Gao K. 2004. Roles of carbonic anhydrase in photosynthesis of Skeletonema costatum.
Journal of Plant Physiology and Molecular Biology. 30(5):511-516.
34. 陈雄文, 高坤山. 2004. 赤潮藻中肋骨条藻的光合作用对海水PH和N变化的响应. 水生生物
学报. 28(6):635-639.
35. 蒋汉明, 高坤山. 2004. 氮源及其浓度对三角褐指藻生长和脂肪酸组成的影响. 水生生物
学报. 28(5):545－551.
36. Gao K and Ye C. 2003. Culture of the terrestrial cyanobacterium, Nostoc flagelliforme
(Cyanophyceae), under aquatic conditions. Journal of Phycology. 39(3):617-623. (IF:2.502)
37. Zou D, Gao K and Xia J. 2003. Photosynthetic utilization of inorganic carbon in the economic
brown alga, Hizikia fusiforme (Sargassaceae) from the South China Sea. Journal of Phycology. 39
(6):1095 -1100. (IF:2.502)
38. Helbling E, Gao K, Goncalves R, Wu H, Villafańe V. 2003. Utilization of solar UV radiation by
coastal phytoplankton assemblages off SE China when exposed to fast mixing. Marine Ecology
Progress Series. 259:59-66. (IF:2.315)
39. Hu H and Gao K. 2003. Optimization of growth and fatty acid composition of a unicellular marine
picoplankton, Nannochloropsis sp., with enriched carbon sources. Biotechnology Letters. 25(5):
421-425. (IF:1.108)
40. Yang Y and Gao K. 2003. Effects of CO2 concentrations on the freshwater microalgae,
Chlamydomonas reinhardtii, Chlorella pyrenoidosa and Scenedesmus obliquus (Chlorophyta).
Journal of Applied Phycology. 15(5):379-389. (IF:0.992)
41. Chen X and Gao K. 2003. Effects of CO2 concentrations on the activity of photosynthetic CO2
fixation and extracellular carbonic anhydrase in the marine diatom Skeletonema costatum. Chinese
Science Bulletin. 48(23): 2616-2620. (IF:0.783)
42. Xia J and Gao K. 2003. Effects of doubled atmospheric CO2 concentration on the photosynthesis
and growth of Chlorella pyrenoidosa cultured at varied levels of light. Fisheries Science. 69(4):767-
771. (IF:0.661)
43. 邹定辉, 高坤山. 2003. 低潮干出状态下潮间带海藻的某些光合生理特性. 植物生理学
通讯. 39(5): 525-530.
44. 蒋汉明, 高坤山. 2003. 两种海洋微藻脂肪酸组成研究. 科学技术与工程. 3(1):46-48.
45. 李运广, 高坤山. 2003. 盐胁迫对地木耳和葛仙米生理生化特性的影响. 水生生物学报.
27(3):227-231.
46. 陈雄文, 高坤山. 2003. CO2浓度对中肋骨条藻的光合无机碳吸收和胞外碳酸酐酶活性
的影响. 科学通报. 48(21):2275-2279.
47. 邹定辉, 高坤山. 2003. 在水生与气生状态下石莼光合作用对光照和温度的响应. 植物
学通报. 20(6):713-722.
联系方式:
联系地址： 广东省汕头市大学路243号，汕头大学海洋生物研究所，515063
联系电话： 0754-82903977
传 真： 0754-82903977
E- mail： ksgao@stu.edu.cn
高坤山 男，1958年9月出生, 教授，博士生导师
主要研究方向：藻类生理生态。1982年毕业于中国海洋大学水产学院， 83年10月作为教育部出国研究生留学日本, 86年3月获东京水产大学硕士学位，89年3月获京都大学农学博士学位。1989年4月至92年7月，任日本关西综合环境中心客座研究员；1992年8月至94年12月，夏威夷大学博士后。1995年1月至1996年12月，汕头大学副教授。1997年入选中国科学院“百人计划”研究员、博士生导师，汕头大学教授，海洋生物研究所所长。
1996年获国家杰出青年科学基金资助，已完成国家自然科学基金重点项目“大气CO2浓度升高与水生生物的关系”、面上项目及国家海洋“863”项目各一项。 目前承担国家自然科学基金重点项目“海洋植物对阳光紫外辐射响应机制的研究”（2005.1-2008.12）。
在20多年的研究历程中，在国内外刊物上表论文90多篇，其中在国际主流刊物发表论文65篇（SCI收录51篇）。多数被国际同行学者在研究论文、综述（reviews)及专著中反复引用（正面引用）。
论文与专著中多次引用。现任〖Journal of Applied Phycology〗（Kluwer Academic Publisher, SCI收录） 唯一中国籍编委 (1998－), 《水生生物学报》、《海洋科学》编委， 国际藻类学会与美国藻类学会终身会员，日本、欧洲藻类学会会员，中国藻类学会理事。国际应用藻类学会执行委员（executive committee, 2005-)。
当前研究重点：
1)海洋生态与光生物学；
2)藻类环境生理,全球环境变化与藻类关系的研究；
3)应用藻类学,经济藻类养殖或培养的生理学基础；
4)藻类光合固碳机理,微藻与海藻类无机碳利用与浓缩机制等。
近年主要论文：
1. Gao K, Wu Y, Li G, Wu H, Villafańe V and Helbling E. 2007. Solar UV-radiation drives CO2-
fixation in marine phytoplankton: A double-edged sword. Plant Physiology. 144:54-59. (IF:6.114)
2. Villafańe V, Gao K, Li P, Li G and Helbling E. 2007. Vertical mixing within the epilimnion
modulates UVR induced photoinhibition in tropical freshwater phytoplankton from southern
China. Freshwater Biology.52:1260-1270. (IF:2.797)
3. Gao K and Ye C. 2007. Photosynthetic insensitivity of Nostoc flagelliforme to solar UV radiation
while rehydrated or desiccated. Journal of Phycology. 43(4):628-635 (IF:2.502)
4. Gao K, Li G, Helbling E and Villafańe V. 2007. Variability of UVR effects on photosynthesis of
summer phytoplankton assemblages from a tropical coastal area of the South China Sea.
Photochemistry and Photobiology. 83:802-809. (IF:2.147)
5. Jiang H, Gao K and Helbling E. 2007. Effects of solar UV radiation on germination of conchospores
and morphogenesis of sporelings inPorphyra haitanensis (Rhodophyta). Marine Biology. 151:1751-
1759. (IF:1.754)
6. Gao K, Guan W and Helbling E. 2007. Effects of solar ultraviolet radiation on photosynthesis of the
marine red tide alga Heterosigma akashiwo (Raphidophyceae). Journal of Photochemistry and
Photobiology B: Biology. 86:140-148. (IF:1.597)
7. Zou D, Gao K, Xia J, Xu Z, Zhang X and Liu S. 2007. responses of dark respiration in the light to
desiccation and temperature in the intertidal macroalga, Ulva lactuca (Chorophyta) during emersion.
Phycologia. 46(3):139-146. (IF:1.271)
8. 徐军田, 高坤山. 2007. 阳光紫外辐射对绿藻石莼光化学效率的影响. 海洋学报. 29(1):129-134.
9. 关万春, 高坤山. 2007. 阳光紫外辐射对两种赤潮藻类光化学效率的影响. 水生生物学报.
31(4):153-158.
10. Zou D, Gao K and Ruan Z. 2006. Seasonal pattern of reproduction of Hizikia fusiformis
(Sargassaceae, Phaeophyta) from Nanao Island, Shantou,China. Journal of Applied Phycology. 18:
195-201. (IF:0.992)
11. Helbling E, Gao K, Ai H, Ma Z and Villafańe V. 2006.Differential responses of Nostoc sphaeroides
and Arthrospira platensis to solar ultraviolet radiation exposure. Journal of Applied Phycology. 18
(1):57-66. (IF:0.992)
12. Ma Z, Gao K and Watanabe T. 2006. Effects of dissolved organic matter on the growth and
pigments synthesis of Spirulina platensis (Arthrospira)*. Progress in Natural Science. 16(1):
50-54. (IF:0.444)
13. Wu H, Gao K, Villafańe V, Watanabe T and Helbling E. 2005. Effect of solar UV radiation on
morphology and photosynthesis of filamentous cyanobacterium Arthrospira platensis. Applied and
Environmental Microbiology. 71(9):5004-5013. (IF:3.818)
14. Villafańe V, Gao K and Helbling E. 2005. Short-and long-term effects of solar ultraviolet radiation
on the red algae Porphyridium cruentum (S. F. Gray) N?geli. Photochemical & Photobiological
Sciences. 4(4):376-382. (IF:2.117)
15. Xia J and Gao K. 2005. Impacts of elevated CO2 concentration on biochemical composition,
carbonic anhydrase, and nitrate reductase activity of freshwater green algae. Journal of Integrative
Plant Biology. 47(6):668-675. (IF:0.432)
16. Zou D and Gao K. 2005. Regulation of gamete release in the economic brown seaweed Hizikia
fusiforme (Phaeophyta). Biotechnology Letters. 27:915-918. (IF:1.108)
17. Zou D and Gao K. 2005. Photosynthetic characteristics of the economic brown seaweed Hizikia
fusiforme (Sargassaceae, Phaeophyta), with special reference to its“leaf” and receptacle.
Journal of Applied Phycology. 17:255-259. (IF:0.992)
18. Wu H, Gao K, Ma Z and Watanabe T. 2005. Effects of solar ultraviolet radiation on biomass
production and pigment contents of Spirulina platensis in commercial operations under sunny and
cloudy weather conditions. Fisheries Science. 71(2):454-456. (IF:0.661)
19. Zou D and Gao K. 2005. Ecophysiological characteristics of four intertidal marine macroalgae
during emersion along Shantou coast of China, with a special reference to the relationship of
photosynthesis and CO2. Acta Oceanologica Sinica. 24(3):105-113. (IF:0.507)
20. Ji Y, Gao K and Tanaka J. 2005. Photosynthetic recovery of desiccated intertidal seaweeds after
rehydration. Progress in Natural Science. 15(8):689-693. (IF:0.444)
21. Wu H, Gao K and Watanabe T. 2005. Effects of solar ultraviolet radiation on the photochemical
efficiency, photosynthetic pigments and biomass production of Spirulina platensis. Progress in
Natural Science. 15(5): 477-480. (IF:0.444)
22. 吴红艳, 高坤山, 渡辺辉夫. 2005. 静止和充气培养条件下短期紫外辐射对钝顶螺旋藻光化
学效率的影响. 水生生物学报.29 (6): 674-677.
23. Gao K and Ai H. 2004. Relationship of growth and photosynthesis with colony size in an edible
cyanobacterium, Ge-Xian-Mi Nostoc (Cyanophyceae). Journal of Phycology. 40(3):523-526.
(IF:2.502)
24. Jiang H and Gao K. 2004. Effects of lowering temperature during culture on the production of
polyunsaturated fatty acids in the marine diatom Phaeodactylum tricornutum (Bacillariophyceae).
Journal of Phycology. 40(4):651-654. (IF:2.502)
25. Chen X and Gao K. 2004. Photosynthetic utilisation of inorganic carbon and its regulation in the
marine diatom Skeletonema costatum. Functional Plant Biology. 31(4):1027-1033. (IF:2.496)
26. Chen X and Gao K. 2004. Characterization of diurnal photosynthetic rhythms in the marine diatom
Skeletonema costatum grown in synchronous culture under ambient and elevated CO2. Functional
Plant Biology. 31(4):399-404. (IF:2.496)
27. Li Y and Gao K. 2004. Photosynthetic physiology and growth as a function of colony size in the
cyanobacterium Nostoc sphaeroides. European Journal of Phycology. 39(1):9-15. (IF:2.064)
28. Qiu B, Zhang A, Liu Z and Gao K. 2004. Studies on the photosynthesis of the terrestrial
cyanobacterium Nostoc flagelliforme subjected to desiccation and subsequent rehydration.
Phycologia. 43(5):521-528. (IF:1.271)
29. Ye C and Gao K. 2004. Photosynthetic response to salt of aquatic-living colonies of the terrestrial
cyanobacterium Nostoc flagelliforme. Journal of Applied Phycology. 16(6):477-481. (IF:0.992)
30. Gao K and Ji Y. 2004. Quantitative evaluation of wind effect during emersion on Porhpyra
haitanensis (Rhodophyta), a farmed species in southern china. Fisheries science. 70(4):710-712.
(IF:0.661)
31. Zou D and Gao K. 2004. Comparative mechanisms of photosynthetic carbon acquisition in Hizikia
fusiforme under submersed and emersed conditions. Acta Botanica Sinica. 46(10):1178-1185.
(IF:0.432)
32. Zou D and Gao K. 2004. Exogenous carbon acquisition of photosynthesis in Porphyra haitanensis
(Bangiales, Rhodophyta) under emersed state. Progress in Natural Science. 14(2):138-144.
(IF:0.444)
33. Chen X and Gao K. 2004. Roles of carbonic anhydrase in photosynthesis of Skeletonema costatum.
Journal of Plant Physiology and Molecular Biology. 30(5):511-516.
34. 陈雄文, 高坤山. 2004. 赤潮藻中肋骨条藻的光合作用对海水PH和N变化的响应. 水生生物
学报. 28(6):635-639.
35. 蒋汉明, 高坤山. 2004. 氮源及其浓度对三角褐指藻生长和脂肪酸组成的影响. 水生生物
学报. 28(5):545－551.
36. Gao K and Ye C. 2003. Culture of the terrestrial cyanobacterium, Nostoc flagelliforme
(Cyanophyceae), under aquatic conditions. Journal of Phycology. 39(3):617-623. (IF:2.502)
37. Zou D, Gao K and Xia J. 2003. Photosynthetic utilization of inorganic carbon in the economic
brown alga, Hizikia fusiforme (Sargassaceae) from the South China Sea. Journal of Phycology. 39
(6):1095 -1100. (IF:2.502)
38. Helbling E, Gao K, Goncalves R, Wu H, Villafańe V. 2003. Utilization of solar UV radiation by
coastal phytoplankton assemblages off SE China when exposed to fast mixing. Marine Ecology
Progress Series. 259:59-66. (IF:2.315)
39. Hu H and Gao K. 2003. Optimization of growth and fatty acid composition of a unicellular marine
picoplankton, Nannochloropsis sp., with enriched carbon sources. Biotechnology Letters. 25(5):
421-425. (IF:1.108)
40. Yang Y and Gao K. 2003. Effects of CO2 concentrations on the freshwater microalgae,
Chlamydomonas reinhardtii, Chlorella pyrenoidosa and Scenedesmus obliquus (Chlorophyta).
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44. 蒋汉明, 高坤山. 2003. 两种海洋微藻脂肪酸组成研究. 科学技术与工程. 3(1):46-48.
45. 李运广, 高坤山. 2003. 盐胁迫对地木耳和葛仙米生理生化特性的影响. 水生生物学报.
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46. 陈雄文, 高坤山. 2003. CO2浓度对中肋骨条藻的光合无机碳吸收和胞外碳酸酐酶活性
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47. 邹定辉, 高坤山. 2003. 在水生与气生状态下石莼光合作用对光照和温度的响应. 植物
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联系方式:
联系地址： 广东省汕头市大学路243号，汕头大学海洋生物研究所，515063
联系电话： 0754-82903977
传 真： 0754-82903977
E- mail： ksgao@stu.edu.cn
高坤山老师相关教学资源
- 汕头大学教学资源
- 高坤山老师课程教学资源
